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The Origin of Species · Chapter

Acclimatisation explained

Darwin explores the capacity of species to adapt to different climates, arguing that acclimatisation is a product of both habit and natural selection.

What happens

Darwin explores the capacity of species to adapt to different climates, arguing that acclimatisation is a product of both habit and natural selection. He presents evidence from plants and animals, including observations by Hooker, Thwaites, and Watson, and discusses the flexibility of constitution common to many organisms, concluding that natural selection of innate variations often overrides the effects of habit.

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Narrative arc

Story tension across the book — this chapter sits in Exposition.

SetupEscalationAftermathClimax
  • Setup
  • Escalation
  • Breaking point
  • Aftermath
  • Closing

This passage marks the story's dramatic climax because Darwin shifts from a defensive posture against objectio…

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Themes in this chapter

Characters to notice

  • Dr. Hooker

    Provided evidence of pines and rhododendrons from different Himalayan altitudes showing varying cold resistance.

  • Mr. H. C. Watson

    Observed acclimatisation in European plants brought from the Azores to England.

  • Charles Darwin

    Author of the chapter, presenting arguments and observations on acclimatisation.

Key passages

  • “Habit is hereditary with plants, as in the period of flowering, in the time of sleep, in the amount of rain requisite for seeds to germinate, etc.”

    Plants inherit habits such as when they flower, when they rest, and how much rain they need to sprout.

    Introduces the concept of hereditary habits in plants.

  • “We have reason to believe that species in a state of nature are closely limited in their ranges by the competition of other organic beings quite as much as, or more than, by adaptation to particular climates.”

    The ranges of wild species are often restricted more by competition with other organisms than by climate alone.

    Highlights the role of biotic interactions in limiting species distribution.

  • “On the whole, we may conclude that habit, or use and disuse, have, in some cases, played a considerable part in the modification of the constitution and structure; but that the effects have often been largely combined with, and sometimes overmastered by, the natural selection of innate variations.”

    Overall, habit and use/disuse sometimes modify an organism's constitution, but natural selection of inborn variations usually plays a larger role.

    Summarises Darwin's conclusion on the relative importance of habit versus natural selection in acclimatisation.